Human remains dating back to the Late Upper Palaeolithic period over 13,000 years ago has revealed a previously unknown "fourth strand" of ancient European ancestry.
This new lineage stems from populations of hunter-gatherers that split from western hunter-gatherers shortly after the 'out of Africa' expansion some 45,000 years ago and went on to settle in the Caucasus region, where southern Russia meets Georgia today.
Here these hunter-gatherers remained for millennia, isolated as the Ice Age culminated in the last 'Glacial Maximum' some 25,000 years ago. They weathered the cold in the relative shelter of the Caucasus mountains until eventual thawing allowed movement and brought them into contact with other populations, likely from further east.